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Date: Time: Abstract The invariant natural killer T iNKT cells are a subset of T lymphocytes that share characteristics of both innate and adaptive immunity. These cells are defined as T cells restricted by the lipid antigen presenting molecule CD1d and express an invariant T cell receptor TCR as well as classical natural killer NK cell markers.

Upon stimulation iNKT cells rapidly produce large amounts of cytokines and chemokines and activate various other cell types including NK cells and dendritic cells DCs. In addition they are involved in the immune responses against tumors and in regulating autoimmune and inflammatory diseases.

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Several viruses have been shown to evade human iNKT cell responses by down- regulating CD1d expression in par söker man i skön cells. The mechanism behind CD1d down-regulation was dependent on inhibition of CD1d recycling, and interaction between CD1d and Vpu in early endosomal compartments. Vpu down-regulates different host-proteins involved in innate immunity.

Interestingly, down-regulation of CD1d and CD4 was found to occur by distinct mechanisms. Commando 3 full movie tfpdl Collectively, these findings suggest that the Vpu protein of HIV-1 may play a significant dejtingsajt frykerud in evasion from CD1d-specific immune responses, and moreover support the notion that iNKT cells play an important role in the host defence against HIV In addition to recognition of exogenous lipid antigens presented by CD1d, iNKT cells can be activated by cytokine or surface receptor stimulation with or without presentation of endogenous lipids.

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These alternative pathways for iNKT cell activation may be particularly important in settings where target cells express no or low levels of CD1d, such as transformed or virus infected cells. In summary, the work presented in this thesis expands our knowledge about how iNKT cells respond to target cells and how viruses work to counteract such recognition.

List of papers: I. Markus Moll, Sofia K. Inhibition of lipid antigen presentation in dendritic cells by HIV-1 Vpu interference with CD1d recycling from endosomal compartments. Institution: Karolinska Institutet. Supervisor: Moll, Markus. This Collection. United States. South Korea.

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